Shopping on line can be easy, simple and save you lots of money. It can also take a lot of your time, frustrate you, and result in unwanted purchases. Now the same can be said for regular high street shopping, but with the vast opportunity presented by the Internet it will pay you to spend a few minutes reading this and understanding how to better optimize your Animals shopping experience:
1. Compare - without doubt the biggest advantage that the Animals offers shoppers today is the ability to compare thousands of Animals at a time. This is a great thing, but not necessarily all the time! Too much can be daunting at times so take advantage of the great comparison sites and where possible let them do the hard work for you.
2. Research - if it has been said it will be on the internet. Ignorance is no longer a justifiable reason for buying the wrong thing. Take the time to research in detail everything that you could possible want to know about
3. Testimonials - don't know anybody that has bought a Animals? Wrong! If the Animals is good the internet will let you know. Use the Internet as a friend and get testimonials before you buy.
4. Questions - Got a question about Animals then search the Forums, FAQ's, Blogs etc. Don't be afraid to ask .....
5. Reputation - Never heard of the company selling Animals? Don't worry, no reason why you should know every company in the world, but you know someone that does! Use the internet to find out what people are saying about Animals and build up a picture of their reputation for sales, returns, customer service, delivery etc.
6. Returns - still worried that even after all of the above your Animals wont be what you want? Check out the returns policy. There is so much competition now that someone, somewhere is bound to offer the terms that you are comfortable with.
7. Feedback - happy with your Animals then let people know, after all you are depending on others people input in your buying decision, so why not give a little back.
8. Security - check for the yellow padlock on the Animals site before you buy, and the s after http:/ /i.e. https:// = a secure site
9. Contact - got a question about Animals, or want to leave a comment then check out the sites contact page. Reputable companies have them and respond.
10. Payment - ready to pay for your Animals, then use your credit card or PayPal! Be aware of companies that don't accept them, there may be genuine reasons but given the huge amount of choice you have when buying online there is no reason at all not to buy via credit card or PayPal.
{{Taxobox| color = pink| name = Animals| fossil_range =
Ediacaran - Recent],
Tawny Owl, Siberian Tiger,
European garden spider,
White-lipped snail, Green Sea Turtle,
Solitary bee, Asian arowana,
Barbary Macaque,
Sawfish and a Marbled White (butterfly).]| unranked_regnum =
Opisthokonta],
Systema Naturae| subdivision_ranks =
Phylum| subdivision =
Subregnum Parazoa
(alternatively)
Calcarea
SilicareaSubregnum Eumetazoa
- Trichoplax
- Radiata (unranked)
- Bilateria (unranked)
-->,
Iciligorgia schrammi, and a sea rod,
Plexaurella nutans.
Animals are a major group of
multicellular organisms, of the
Kingdom (biology) Animalia or
Metazoa. Their
body plan becomes fixed as they develop, usually early on in their
ontogeny as embryos, although some undergo a process of
metamorphosis (biology) later on in their life. Most animals are motile - can move spontaneously and independently.
Etymology
The word "animal" comes from the
Latin word
animal, of which
animalia is the plural, and is derived from
anima, meaning vital breath or soul. In everyday colloquial usage, the word usually refers to non-
human animals. The biological definition of the word refers to all members of the Kingdom Animalia. Therefore, when the word "animal" is used in a biological context, humans are included.
Characteristics
Animals have several characteristics that set them apart from other living things. Animals are
eukaryote and usually
multicellular (although see
Myxozoa), which separates them from
bacteria and most protists. They are
heterotrophic, generally digesting food in an internal chamber, which separates them from
plants and algae. They are also distinguished from plants, algae, and
fungus by lacking
cell walls. All animals are motile, if only at certain life stages. Embryos pass through a Blastula, which is a characteristic exclusive to animals.
Structure
With a few exceptions, most notably the
sponges (Phylum Porifera), animals have bodies differentiated into separate biological tissue. These include
muscles, which are able to contract and control locomotion, and
nervous system, which sends and processes signals. There is also typically an internal digestion chamber, with one or two openings. Animals with this sort of organization are called metazoans, or
eumetazoans when the former is used for animals in general.
All animals have
eukaryotic cells, surrounded by a characteristic extracellular matrix composed of
collagen and elastic
glycoproteins. This may be calcified to form structures like
animal shells,
bones, and
spicules. During development it forms a relatively flexible framework upon which cells can move about and be reorganized, making complex structures possible. In contrast, other multicellular organisms like plants and fungi have cells held in place by cell walls, and so develop by progressive growth. Also, unique to animal cells are the following intercellular junctions:
tight junctions,
gap junctions, and
desmosomes.
Reproduction and development
lung cell (biology)
staining (biology) with fluorescent dyes undergoing
mitosis, specifically early anaphase.Nearly all animals undergo some form of sexual reproduction. Adults are diploid or
polyploid. They have a few specialized reproductive cells, which undergo meiosis to produce smaller motile
spermatozoon or larger non-motile ovum. These fuse to form
zygotes, which develop into new individuals.
Many animals are also capable of
asexual reproduction. This may take place through parthenogenesis, where fertile eggs are produced without mating, or in some cases through
fragmentation (biology).
A
zygote initially develops into a hollow sphere, called a
blastula, which undergoes rearrangement and differentiation. In sponges, blastula larvae swim to a new location and develop into a new sponge. In most other groups, the blastula undergoes more complicated rearrangement. It first invagination to form a gastrula with a digestive chamber, and two separate
germ layers - an external ectoderm and an internal endoderm. In most cases, a mesoderm also develops between them. These germ layers then differentiate to form tissues and organs.
Most animals grow by indirectly using the energy of sunlight. Plants use this energy to convert sunlight into simple
sugars using a process known as photosynthesis. Starting with the molecules carbon dioxide (CO2) and water (H2O), photosynthesis converts the energy of sunlight into chemical energy stored in the bonds of glucose (C6H12O6) and releases
oxygen (O2). These sugars are then used as the building blocks which allow the plant to grow. When animals eat these plants (or eat other animals which have eaten plants), the sugars produced by the plant are used by the animal. They are either used directly to help the animal grow, or broken down, releasing stored solar energy, and giving the animal the energy required for motion. This process is known as
glycolysis.
Animals who live close to
hydrothermal vents and
cold seeps on the ocean floor are not dependent on the energy of sunlight. Instead, chemosynthesis
archaea and
eubacteria form the base of the food chain.
Origin and fossil record
Animals are generally considered to have
Evolution from a flagellated eukaryote. Their closest known living relatives are the choanoflagellates, collared flagellates that have a morphology similar to the choanocytes of certain sponges.
Molecular studies place animals in a supergroup called the
opisthokonts, which also include the choanoflagellates, fungus and a few small parasitic
protists. The name comes from the posterior location of the flagellum in motile cells, such as most animal spermatozoa, whereas other
eukaryotes tend to have anterior flagella.
The first fossils that might represent animals appear towards the end of the Precambrian, around 575 million years ago, and are known as the
Ediacaran biota. These are difficult to relate to later fossils, however. Some may represent precursors of modern phyla, but they may be separate groups, and it is possible they are not really animals at all. Aside from them, most known animal phyla make a more or less simultaneous appearance during the
Cambrian period, about 542 million years ago. It is still disputed whether this event, called the
Cambrian explosion, represents a rapid divergence between different groups or a change in conditions that made fossilization possible.
Groups of animals
The sponges (Porifera) diverged from other animals early. As mentioned above, they lack the complex organization found in most other phyla. Their cells are differentiated, but in most cases not organized into distinct tissues. Sponges are sessile and typically feed by drawing in water through pores.
Archaeocyatha, which have fused skeletons, may represent sponges or a separate phylum.
Among the eumetazoan phyla, two are radially symmetric and have digestive chambers with a single opening, which serves as both the mouth and the anus. These are the Cnidaria, which include sea anemones, corals, and
jellyfish, and the
Ctenophora or comb jellies. Both have distinct tissues, but they are not organized into organ (anatomy). There are only two main germ layers, the ectoderm and endoderm, with only scattered cells between them. As such, these animals are sometimes called
diploblastic. The tiny Placozoans are similar, but they do not have a permanent digestive chamber.
The remaining animals form a monophyletic group called the
Bilateria. For the most part, they are bilaterally symmetric, and often have a specialized head with feeding and sensory organs. The body is triploblastic, i.e. all three germ layers are well-developed, and tissues form distinct organs. The digestive chamber has two openings, a mouth and an anus, and there is also an internal body cavity called a coelom or pseudocoelom. There are exceptions to each of these characteristics, however - for instance adult echinoderms are radially symmetric, and certain parasitic worms have extremely simplified body structures.
Genetic studies have considerably changed our understanding of the relationships within the Bilateria. Most appear to belong to four major lineages:
Deuterostomes
Ecdysozoa
Platyzoa
Lophotrochozoa
In addition to these, there are a few small groups of bilaterians with relatively similar structure that appear to have diverged before these major groups. These include the Acoelomorpha, Rhombozoa, and Orthonectida. The
Myxozoa, single-celled parasites that were originally considered Protozoa, are now believed to have developed from the Bilateria as well.
Deuterostomes
,
Malurus cyaneusDeuterostomes differ from the other Bilateria, called
protostomes, in several ways. In both cases there is a complete digestive tract. However, in protostomes the initial opening (the
archenteron) develops into the mouth, and an anus forms separately. In deuterostomes this is reversed. In most protostomes cells simply fill in the interior of the gastrula to form the mesoderm, called schizocoelous development, but in deuterostomes it forms through invagination of the endoderm, called enterocoelic pouching. Deuterostomes also have a dorsal, rather than a ventral, nerve chord and their embryos undergo different cleavage.
All this suggests the deuterostomes and protostomes are separate, monophyletic lineages. The main phyla of deuterostomes are the Echinodermata and
Chordate. The former are radially symmetric and exclusively marine, such as
starfish, sea urchins, and
sea cucumbers. The latter are dominated by the
vertebrates, animals with backbones. These include
fish,
amphibians,
reptiles, birds, and mammals.
In addition to these, the deuterostomes also include the
Hemichordata or acorn worms. Although they are not especially prominent today, the important fossil
graptolites may belong to this group.
The Chaetognatha or arrow worms may also be deuterostomes, but more recent studies suggest protostome affinities.
Ecdysozoa
,
Sympetrum flaveolumThe Ecdysozoa are protostomes, named after the common trait of growth by moulting or
ecdysis. The largest animal phylum belongs here, the
Arthropoda, including
insects, spiders, crabs, and their kin. All these organisms have a body divided into repeating segments, typically with paired appendages. Two smaller phyla, the Onychophora and
Tardigrada, are close relatives of the arthropods and share these traits.
The ecdysozoans also include the Nematoda or roundworms, the second largest animal phylum. Roundworms are typically microscopic, and occur in nearly every environment where there is water. A number are important parasites. Smaller phyla related to them are the Nematomorpha or horsehair worms, which are invisible to the unaided eye, and the
Kinorhyncha, Priapulida, and Loricifera. These groups have a reduced coelom, called a pseudocoelom.
The remaining two groups of protostomes are sometimes grouped together as the
Spiralia, since in both embryos develop with spiral cleavage.
Platyzoa
The Platyzoa include the phylum Platyhelminthes, the flatworms. These were originally considered some of the most primitive Bilateria, but it now appears they developed from more complex ancestors.
A number of parasites are included in this group, such as the
flukes and
tapeworms. Flatworms lack a coelom, as do their closest relatives, the microscopic
Gastrotricha.
The other platyzoan phyla are microscopic and pseudocoelomate. The most prominent are the
Rotifera or rotifers, which are common in aqueous environments. They also include the
Acanthocephala or spiny-headed worms, the Gnathostomulida,
Micrognathozoa, and possibly the
Cycliophora. These groups share the presence of complex jaws, from which they are called the Gnathifera.
Lophotrochozoa
The
Lophotrochozoa include two of the most successful animal phyla, the
Mollusca and Annelida. The former includes animals such as snails, clams, and
squids, and the latter comprises the segmented worms, such as
earthworms and leeches. These two groups have long been considered close relatives because of the common presence of trochophore larvae, but the annelids were considered closer to the arthropods, because they are both segmented. Now this is generally considered convergent evolution, owing to many morphological and genetic differences between the two phyla.
The Lophotrochozoa also include the Nemertea or ribbon worms, the
Sipuncula, and several phyla that have a fan of cilia around the mouth, called a lophophore. These were traditionally grouped together as the lophophorates, but it now appears they are paraphyletic, some closer to the Nemertea and some to the Mollusca and Annelida. They include the
Brachiopoda or lamp shells, which are prominent in the fossil record, the
Entoprocta, the Phoronida, and possibly the Bryozoa or moss animals.
Model organisms
Because of the great diversity found in animals, it is more economical for scientists around the world concert their efforts on a small number of chosen species so that connections can be drawn from their work and conclusions extrapolated about how animals function in general. Because they are easy to keep and breed, the fruit fly
Drosophila melanogaster and the nematode
Caenorhabditis elegans have long been the most intensively studied metazoan model organism, and among the first lifeforms to be genetically sequenced. This was facilitated by the severely reduced state of their genomes, but the double-edged sword here is that with many
genes,
introns and genetic linkage lost, these ecdysozoans can teach us little about the origins of animals in general. The extent of this type of evolution within the superphylum will be revealed by the crustacean, annelid, and molluscan
genome projects currently in progress. Analysis of the starlet sea anemone genome has emphasised the importance of sponges, placozoans, and choanoflagellates, also being sequenced, in explaining the arrival of 1500 ancestral genes unique to the Eumetazoa.
An analyse of the homoscleromorph sponge Oscarella carmela also suggests that the last common ancestor of sponges and the eumetazoan animals were more comlex than previously assumed. Mitochondrial Genome of the Homoscleromorph Oscarella carmela (Porifera, Demospongiae) Reveals Unexpected Complexity in the Common Ancestor of Sponges and Other Animals
Oxford JournalsOther model organisms belonging to the animal family include mouse (Mus musculus) and zebrafish (
Danio rerio}.
History of classification
, known as the "father of modern
taxonomy".
Aristotle divided the living world between animals and
plants, and this was followed by Carolus Linnaeus in the first hierarchical classification. Since then biologists have begun emphasizing evolutionary relationships, and so these groups have been restricted somewhat. For instance, microscopic
protozoa were originally considered animals because they move, but are now treated separately.
In
Carolus Linnaeus original scheme, the animals were one of three kingdoms, divided into the classes of
Vermes,
Insecta, Fish, Amphibia,
bird, and
Mammalia. Since then the last four have all been subsumed into a single phylum, the
chordate, whereas the various other forms have been separated out. The above lists represent our current understanding of the group, though there is some variation from source to source.
See also
Notes
References
- Klaus Nielsen. Animal Evolution: Interrelationships of the Living Phyla (2nd edition). Oxford Univ. Press, 2001.
- Knut Schmidt-Nielsen. Animal Physiology: Adaptation and Environment. (5th edition). Cambridge Univ. Press, 1997.
External links
- Tree of Life Project
- Animal Diversity Web - University of Michigan's database of animals, showing taxonomic classification, images, and other information.
- ARKive - multimedia database of worldwide endangered/protected species and common species of UK.
- Scientific American Magazine (December 2005 Issue) - Getting a Leg Up on Land About the evolution of four-limbed animals from fish.
{{Taxobox| color = pink| name = Animals| fossil_range =
Ediacaran - Recent],
Tawny Owl, Siberian Tiger, European garden spider,
White-lipped snail, Green Sea Turtle, Solitary bee,
Asian arowana,
Barbary Macaque,
Sawfish and a Marbled White (butterfly).]| unranked_regnum =
Opisthokonta],
Systema Naturae| subdivision_ranks =
Phylum| subdivision =
Subregnum Parazoa
(alternatively)
CalcareaSilicarea
Subregnum Eumetazoa
- Trichoplax
- Radiata (unranked)
- Bilateria (unranked)
- Orthonectida
- Rhombozoa
- Acoelomorpha
- Chaetognatha
- Superphylum Deuterostome
- Chordate
- Hemichordata
- Echinodermata
- Xenoturbellida
- Vetulicolia extinction
- Superphylum Ecdysozoa
- Superphylum Platyzoa
- Superphylum Lophotrochozoa
-->,
Iciligorgia schrammi, and a sea rod,
Plexaurella nutans.
Animals are a major group of
multicellular organisms, of the
Kingdom (biology) Animalia or
Metazoa. Their
body plan becomes fixed as they develop, usually early on in their ontogeny as
embryos, although some undergo a process of metamorphosis (biology) later on in their life. Most animals are
motile - can move spontaneously and independently.
Etymology
The word "animal" comes from the Latin word
animal, of which
animalia is the plural, and is derived from
anima, meaning vital breath or soul. In everyday colloquial usage, the word usually refers to non-human animals. The biological definition of the word refers to all members of the Kingdom Animalia. Therefore, when the word "animal" is used in a biological context, humans are included.
Characteristics
Animals have several characteristics that set them apart from other living things. Animals are eukaryote and usually multicellular (although see Myxozoa), which separates them from bacteria and most
protists. They are heterotrophic, generally digesting food in an internal chamber, which separates them from plants and
algae. They are also distinguished from plants, algae, and fungus by lacking cell walls. All animals are motile, if only at certain life stages. Embryos pass through a Blastula, which is a characteristic exclusive to animals.
Structure
With a few exceptions, most notably the
sponges (Phylum Porifera), animals have bodies differentiated into separate
biological tissue. These include muscles, which are able to contract and control locomotion, and nervous system, which sends and processes signals. There is also typically an internal
digestion chamber, with one or two openings. Animals with this sort of organization are called metazoans, or eumetazoans when the former is used for animals in general.
All animals have eukaryotic cells, surrounded by a characteristic extracellular matrix composed of collagen and elastic
glycoproteins. This may be calcified to form structures like
animal shells,
bones, and spicules. During development it forms a relatively flexible framework upon which cells can move about and be reorganized, making complex structures possible. In contrast, other multicellular organisms like plants and fungi have cells held in place by cell walls, and so develop by progressive growth. Also, unique to animal cells are the following intercellular junctions: tight junctions,
gap junctions, and desmosomes.
Reproduction and development
lung cell (biology) staining (biology) with fluorescent
dyes undergoing
mitosis, specifically early anaphase.Nearly all animals undergo some form of sexual reproduction. Adults are diploid or
polyploid. They have a few specialized reproductive cells, which undergo
meiosis to produce smaller motile
spermatozoon or larger non-motile ovum. These fuse to form
zygotes, which develop into new individuals.
Many animals are also capable of asexual reproduction. This may take place through
parthenogenesis, where fertile eggs are produced without mating, or in some cases through
fragmentation (biology).
A
zygote initially develops into a hollow sphere, called a blastula, which undergoes rearrangement and differentiation. In sponges, blastula larvae swim to a new location and develop into a new sponge. In most other groups, the blastula undergoes more complicated rearrangement. It first
invagination to form a gastrula with a digestive chamber, and two separate germ layers - an external ectoderm and an internal endoderm. In most cases, a mesoderm also develops between them. These germ layers then differentiate to form tissues and organs.
Most animals grow by indirectly using the energy of
sunlight. Plants use this energy to convert sunlight into simple sugars using a process known as
photosynthesis. Starting with the molecules carbon dioxide (CO2) and water (H2O), photosynthesis converts the energy of sunlight into chemical energy stored in the bonds of
glucose (C6H12O6) and releases
oxygen (O2). These sugars are then used as the building blocks which allow the plant to grow. When animals eat these plants (or eat other animals which have eaten plants), the sugars produced by the plant are used by the animal. They are either used directly to help the animal grow, or broken down, releasing stored solar energy, and giving the animal the energy required for motion. This process is known as
glycolysis.
Animals who live close to hydrothermal vents and cold seeps on the ocean floor are not dependent on the energy of sunlight. Instead, chemosynthesis
archaea and
eubacteria form the base of the food chain.
Origin and fossil record
Animals are generally considered to have Evolution from a flagellated eukaryote. Their closest known living relatives are the
choanoflagellates, collared flagellates that have a morphology similar to the choanocytes of certain sponges.
Molecular studies place animals in a supergroup called the opisthokonts, which also include the choanoflagellates,
fungus and a few small parasitic
protists. The name comes from the posterior location of the flagellum in motile cells, such as most animal spermatozoa, whereas other
eukaryotes tend to have anterior flagella.
The first fossils that might represent animals appear towards the end of the
Precambrian, around 575 million years ago, and are known as the Ediacaran biota. These are difficult to relate to later fossils, however. Some may represent precursors of modern phyla, but they may be separate groups, and it is possible they are not really animals at all. Aside from them, most known animal phyla make a more or less simultaneous appearance during the Cambrian period, about 542 million years ago. It is still disputed whether this event, called the Cambrian explosion, represents a rapid divergence between different groups or a change in conditions that made fossilization possible.
Groups of animals
The sponges (Porifera) diverged from other animals early. As mentioned above, they lack the complex organization found in most other phyla. Their cells are differentiated, but in most cases not organized into distinct tissues. Sponges are sessile and typically feed by drawing in water through pores. Archaeocyatha, which have fused skeletons, may represent sponges or a separate phylum.
Among the eumetazoan phyla, two are radially symmetric and have digestive chambers with a single opening, which serves as both the mouth and the anus. These are the
Cnidaria, which include sea anemones, corals, and jellyfish, and the
Ctenophora or comb jellies. Both have distinct tissues, but they are not organized into
organ (anatomy). There are only two main germ layers, the ectoderm and endoderm, with only scattered cells between them. As such, these animals are sometimes called diploblastic. The tiny
Placozoans are similar, but they do not have a permanent digestive chamber.
The remaining animals form a
monophyletic group called the
Bilateria. For the most part, they are bilaterally symmetric, and often have a specialized head with feeding and sensory organs. The body is
triploblastic, i.e. all three germ layers are well-developed, and tissues form distinct organs. The digestive chamber has two openings, a mouth and an anus, and there is also an internal body cavity called a coelom or pseudocoelom. There are exceptions to each of these characteristics, however - for instance adult echinoderms are radially symmetric, and certain parasitic worms have extremely simplified body structures.
Genetic studies have considerably changed our understanding of the relationships within the Bilateria. Most appear to belong to four major lineages:
Deuterostomes
Ecdysozoa
Platyzoa
Lophotrochozoa
In addition to these, there are a few small groups of bilaterians with relatively similar structure that appear to have diverged before these major groups. These include the
Acoelomorpha,
Rhombozoa, and
Orthonectida. The Myxozoa, single-celled parasites that were originally considered Protozoa, are now believed to have developed from the Bilateria as well.
Deuterostomes
,
Malurus cyaneusDeuterostomes differ from the other Bilateria, called protostomes, in several ways. In both cases there is a complete digestive tract. However, in protostomes the initial opening (the
archenteron) develops into the mouth, and an anus forms separately. In deuterostomes this is reversed. In most protostomes cells simply fill in the interior of the gastrula to form the mesoderm, called schizocoelous development, but in deuterostomes it forms through
invagination of the endoderm, called enterocoelic pouching. Deuterostomes also have a dorsal, rather than a ventral, nerve chord and their embryos undergo different cleavage.
All this suggests the deuterostomes and protostomes are separate, monophyletic lineages. The main phyla of deuterostomes are the Echinodermata and Chordate. The former are radially symmetric and exclusively marine, such as
starfish, sea urchins, and
sea cucumbers. The latter are dominated by the vertebrates, animals with backbones. These include
fish, amphibians,
reptiles, birds, and
mammals.
In addition to these, the deuterostomes also include the Hemichordata or acorn worms. Although they are not especially prominent today, the important fossil graptolites may belong to this group.
The
Chaetognatha or arrow worms may also be deuterostomes, but more recent studies suggest protostome affinities.
Ecdysozoa
,
Sympetrum flaveolumThe Ecdysozoa are protostomes, named after the common trait of growth by moulting or
ecdysis. The largest animal phylum belongs here, the
Arthropoda, including
insects,
spiders,
crabs, and their kin. All these organisms have a body divided into repeating segments, typically with paired appendages. Two smaller phyla, the
Onychophora and
Tardigrada, are close relatives of the arthropods and share these traits.
The ecdysozoans also include the Nematoda or roundworms, the second largest animal phylum. Roundworms are typically microscopic, and occur in nearly every environment where there is water. A number are important parasites. Smaller phyla related to them are the Nematomorpha or horsehair worms, which are invisible to the unaided eye, and the Kinorhyncha, Priapulida, and
Loricifera. These groups have a reduced coelom, called a pseudocoelom.
The remaining two groups of protostomes are sometimes grouped together as the Spiralia, since in both embryos develop with spiral cleavage.
Platyzoa
The
Platyzoa include the phylum Platyhelminthes, the flatworms. These were originally considered some of the most primitive Bilateria, but it now appears they developed from more complex ancestors.
A number of parasites are included in this group, such as the
flukes and
tapeworms. Flatworms lack a coelom, as do their closest relatives, the microscopic
Gastrotricha.
The other platyzoan phyla are microscopic and pseudocoelomate. The most prominent are the
Rotifera or rotifers, which are common in aqueous environments. They also include the Acanthocephala or spiny-headed worms, the
Gnathostomulida,
Micrognathozoa, and possibly the Cycliophora. These groups share the presence of complex jaws, from which they are called the Gnathifera.
Lophotrochozoa
The
Lophotrochozoa include two of the most successful animal phyla, the Mollusca and
Annelida. The former includes animals such as
snails, clams, and
squids, and the latter comprises the segmented worms, such as
earthworms and leeches. These two groups have long been considered close relatives because of the common presence of trochophore larvae, but the annelids were considered closer to the arthropods, because they are both segmented. Now this is generally considered
convergent evolution, owing to many morphological and genetic differences between the two phyla.
The Lophotrochozoa also include the Nemertea or ribbon worms, the
Sipuncula, and several phyla that have a fan of cilia around the mouth, called a lophophore. These were traditionally grouped together as the lophophorates, but it now appears they are
paraphyletic, some closer to the Nemertea and some to the Mollusca and Annelida. They include the Brachiopoda or lamp shells, which are prominent in the fossil record, the
Entoprocta, the Phoronida, and possibly the Bryozoa or moss animals.
Model organisms
Because of the great diversity found in animals, it is more economical for scientists around the world concert their efforts on a small number of chosen species so that connections can be drawn from their work and conclusions extrapolated about how animals function in general. Because they are easy to keep and breed, the fruit fly
Drosophila melanogaster and the nematode
Caenorhabditis elegans have long been the most intensively studied metazoan model organism, and among the first lifeforms to be genetically sequenced. This was facilitated by the severely reduced state of their
genomes, but the double-edged sword here is that with many
genes,
introns and genetic linkage lost, these ecdysozoans can teach us little about the origins of animals in general. The extent of this type of evolution within the superphylum will be revealed by the crustacean, annelid, and molluscan genome projects currently in progress. Analysis of the
starlet sea anemone genome has emphasised the importance of sponges, placozoans, and choanoflagellates, also being sequenced, in explaining the arrival of 1500 ancestral genes unique to the Eumetazoa.
An analyse of the homoscleromorph sponge Oscarella carmela also suggests that the last common ancestor of sponges and the eumetazoan animals were more comlex than previously assumed. Mitochondrial Genome of the Homoscleromorph Oscarella carmela (Porifera, Demospongiae) Reveals Unexpected Complexity in the Common Ancestor of Sponges and Other Animals
Oxford JournalsOther model organisms belonging to the animal family include mouse (Mus musculus) and zebrafish (
Danio rerio}.
History of classification
, known as the "father of modern taxonomy".
Aristotle divided the living world between animals and plants, and this was followed by Carolus Linnaeus in the first hierarchical classification. Since then biologists have begun emphasizing evolutionary relationships, and so these groups have been restricted somewhat. For instance, microscopic protozoa were originally considered animals because they move, but are now treated separately.
In Carolus Linnaeus original scheme, the animals were one of three kingdoms, divided into the classes of
Vermes, Insecta, Fish,
Amphibia,
bird, and
Mammalia. Since then the last four have all been subsumed into a single phylum, the chordate, whereas the various other forms have been separated out. The above lists represent our current understanding of the group, though there is some variation from source to source.
See also
Notes
References
- Klaus Nielsen. Animal Evolution: Interrelationships of the Living Phyla (2nd edition). Oxford Univ. Press, 2001.
- Knut Schmidt-Nielsen. Animal Physiology: Adaptation and Environment. (5th edition). Cambridge Univ. Press, 1997.
External links
- Tree of Life Project
- Animal Diversity Web - University of Michigan's database of animals, showing taxonomic classification, images, and other information.
- ARKive - multimedia database of worldwide endangered/protected species and common species of UK.
- Scientific American Magazine (December 2005 Issue) - Getting a Leg Up on Land About the evolution of four-limbed animals from fish.
Animal
Animal - Clothing, Watches, Footwear, Eyewear, Luggage, Accessories
Animal
BBC - Science & Nature - Animals
A site for anyone interested in animals and the natural world around them. Includes special features, fact sheets, and online quizzes.
CBBC Newsround | Animals
CBBC Newsround - Your stories, your world - first! ... Vote. TV animals Shaun the Sheep, Scooby-Doo or something else? Tell us your fave...
Animal Farming - Vegan Society
Information on how animals are exploited for food and other products and on how to care for vegan companion animals
The Animal Rights FAQ
The Animal Rights FAQ (framed version) ... view the navigation map. U.S and U.K. organisations . view the FAQ by topics
Animal Facts
undefined ... Welcome to our animal fact sheet page! Click on one of the animal types on the menu to the right to get a list of useful animal fact sheets by species.
BBC - Cult - Classic TV - Animal Magic
Animal Magic - Johnny Morris is going to the zoo and you can come too. With Classic clips from the BBC's archives - part of the BBC Cult Website
animals on the underground
A site dedicated to "The Animals" that can be found on London tube maps using a little imagination.
The Learning Zone: Animals
Animals in the learning zone ... Welcome to the animals homepage in the learning zone! The study of animals, their environment and their behaviour is called zoology.